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I've visited two of Carrie Hale litters near Seattle, a each had about 1/4 fluffies I think. It's a Mendelian recessive.
I wonder if fluffy heterozygotes (carriers) have somewhat plusher coats and are selected for -- this might explain the persistence if the allele. It's supposedly not such a good outdoor coat-- more mud, wet, burrs, combing, shedding -- maybe so, but it doesn't look much unlike a border collie or shelty coat to me.
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